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Hedderson TA, Murray DJ, Cox CJ, Nowell TL. Phylogenetic Relationships of Haplolepideous Mosses (Dicranidae) Inferred from rps4 Gene Sequences. Systematic Botany. 2004;29(1):29 - 41. doi:10.1600/036364404772973960
Hedderson TA, Cox CJ, J. Gibbings G. Phylogenetic Relationships of the Wardiaceae (Musci); Evidence from 18s rRNA and rps4 Gene Sequences. The Bryologist. 1999;102(1):26. doi:10.2307/3244455